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Annie O'Malley's dream of owning a secondhand bookstore in tight-knit Audubon Springs is coming true. Still reeling from her parents' sudden deaths, she believes that signs or messages can be found within the pages of books. She loves her job and has made plenty of friends but longs for the acceptance only love and a family can provide.
Tony Iuliano is there to fix the wiring in her shop. He doesn't believe in fate, magic, or the power of a good book. All this combined with his sexy-as-sin presence and all-knowing attitude infuriates Annie. The quirky bookworm intrigues him enough to accept her bet to read a book from her shop. When he discovers a stack of fifty-year-old love letters in the book, he wants to leave the past alone, but seeing it as a sign, Annie challenges him to read the letters and search for the long-lost lovers or lose the bet.
While shaking the dust off several ill-fated romances and fighting their attraction at every turn, the pair soon become the town gossip daily special. If they can stop arguing long enough, they may just discover this particular sign was meant for them all along.